A father recounts to his children - through a series of flashbacks - the journey he and his four best friends took leading up to him meeting their mother.
On their way to Long Island for their wedding weekend, Robin and Barney come across a startling family discovery. Meanwhile, Marshall sees something online that changes the course of his journey east, while Lily confronts Ted about letting go of Robin.
When James reveals that he is getting divorced, Robin worries about how Barney will take the news. Meanwhile, Marshall races to find a way to get to New York in time for the wedding, and Ted deals with being alone at a romantic hotel.
When Lily discovers Ted’s list of things he wants to do in New York before he moves for Chicago, she becomes frustrated that she is the only person who knows about his plans. Meanwhile, Robin and Barney realize they have very little time together before their relatives arrive for their wedding.
While Barney works through his anger at Ted after finding out he still has feelings for Robin, Marshall Skypes in to act as a judge and help settle their dispute. Meanwhile, Lily realizes she is Robin’s only girlfriend.
Barney is upset after Robin wins James's wedding ring during the poker game, and Ted argues with Marshall and Lily over whether he gave them a wedding gift.
When Ted finds himself with three prospects to be his date for the wedding weekend, he chooses poorly. Meanwhile, Barney and Robin have a confrontation with their minister, and Marshall learns more about his driving companion, Daphne.
When Daphne sends a troubling text message to Lily, Marshall enlists the gang's help to remove the message while enforcing the "No Questions Asked" rule.
When Robin and Loretta's conflict escalates, Barney is caught in the middle. Meanwhile, Marshall and Daphne deal with a stowaway on their road trip, and Ted and Cassie try to enjoy a trip to a lighthouse.
Robin and Ted reminisce about past relationships as they search for a drunken Barney on the day of the wedding; Lily and Marshall come to a resolution; Barney befriends two men who are eager to meet women.
Robin and Lily have a fight on the morning of the wedding; Barney can't decide what suit to wear; the gang watches a movie that Ted didn't want them to see.
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